Andrew Joseph Kerkhoff Departments of Biology and Mathematics Kenyon College Gambier, OH 43022
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Andrew Joseph Kerkhoff
Departments of Biology and Mathematics
Kenyon College
Gambier, OH 43022
Phone: 740.427.5734
FAX: 740.427.5741
Education
Ph. D., Biology, 2002. University of New Mexico, Albuquerque, NM.
M.S., Biology, 1997. University of New Mexico, Albuquerque, NM.
B.A., cum laude, English, 1990. Rutgers College, New Brunswick, NJ.
Professional Positions
Assistant Professor, 2005-present, Kenyon College
Postdoctoral Research Associate, 2002-2005, University of Arizona
Undergraduate Advisor, 2002, University of New Mexico
Research Assistant, 1995–1999, University of New Mexico
Teaching And Mentoring Experience
Instructor, 2005-present, Kenyon College:
Introduction to Experimental Biology (Biol109-110)
Elements of Statistics (Math106)
Ecology (Biol228)
Ecology Lab (Biol229)
Global Ecology and Biogeography (Biol328)
Research in Biology (Biol 385/386)
Biological Scaling: Why Size Matters (Biol107 – nonmajors)
Mathematical Biology (Math 258)
Honors Thesis Advisor, 2006-present, Kenyon College
Ellen Thompson, 2008, Biology. A macroecological analysis of tree –
Lepidoptera interactions.
Alexandra Lodge, 2007, Biology. Environmental filtering and limiting similarity
at local and regional scales in eastern deciduous forests.
Kristen Kvernland, 2007, International Studies. On the verge of collapse?
Human and environmental concerns and deforestation in Madagascar.
Guest Lecturer, 2002-2005, University of Arizona:
Core Ecology Seminar (graduate)
Evolution of Plant Form, Function, and Diversity (graduate/undergraduate)
Scaling in Biology (graduate/undergraduate)
Research Supervisor, 2002 – 2005, University of Arizona
Mentor, 2002, Ecological Society of America annual meeting SEEDS Program,
Tucson, AZ
Instructor, 2001, University of New MexicoMind and Nature: The Construction of Ecological Knowledge
Guest Lecturer, 1997-2000, University of New Mexico
Principles of Ecology (undergraduate)
Population Biology (graduate)
Ecological Complexity Seminar (graduate/undergraduate
Teaching Assistant 1996-2001, University of New Mexico
Principles of Ecology
Biology lab for non-majors
Principles of Biology lab
Courses and Workshops
Gordon Research Conference: The Metabolic Basis of Ecology, July 2008.
Lewiston, ME Organizers: Robert Sterner and Mike Kaspari.
Gordon Research Conference: The Metabolic Basis of Ecology and Evolution,
July 2006. Lewiston, ME Organizers: Pablo Marquet and Val Smith.
OSU Mathematical Biosciences Institute Workshop: Global Ecology, June 2006.
Columbus, OH Organizers: John Pastor, John Harte, and David Schimel.
Woodstoich 2004, August 2004, Finse, Norway. Workshop sponsored by the
Norwegian Center for Advanced Study. Organizers: James Elser and Dag
Hessen.
Gordon Research Conference: The Metabolic Basis of Ecology, July 2004.
Lewiston, ME Organizers: James Gillooly and James Brown.
Resilience and Change in Ecosystems, October 2003, Santa Fe, NM. Workshop
cosponsored by the Santa Fe Institute and the Resiliance Alliance.
Organizers: Colleen Webb, Brian Walker, Simon Levin, Erica Jen, and Lars
Hedin
Inquiry methods to Achieve Active Learning in Ecology, August 2003, Savannah,
GA. Workshop sponsored by the Ecological Society of America Education
Section. Instructor: Diane Ebert-May
Innovative Teaching to Achieve Active Learning in Ecology, August 2002,
Tucson, AZ. Workshop sponsored by the Ecological Society of America
Education Section. Instructors: Diane Ebert-May and George Middendorf
Research Grants
NSF Interdisciplinary Training for Undergraduates in Biological and Mathematical
Sciences (UBM) grant. Investigating the Mathematical Biology of Metabolic
Scaling using Manduca InSTaRs (Interdisciplinary Science Training and
Research). Kenyon College 2008 – 2011. ($236,410, PI, co-PIs Chistopher
Gillen, Bradley Hartlaub, Judy Holdener, and Haruhiko Itagaki)
NSF Course Curriculum and Laboratory Improvement Grant, Development and
Dissemination of Computational Science Educational Materials and Curricula
at the Undergraduate Level. Grant to Capitol University 2007 – 2008.
($480,000, $10,000 subcontract, Senior Personnel, Kenyon co-PI Sheryl
Hemkin)
National Parks Service Research Grant, Assessing Floristic Communities along
an Elevational Gradient In Tropical Mexico – Comparison to FlorrisantNational Monument, CO. University of Arizona, 2003 – 2004 ($12,000, Co-PI
with B. Boyle and B. Enquist)
NSF Doctoral Dissertation Improvement Grant, Constraints and drivers of piñon-
juniper woodland dynamics, UNM, 2000 ($6,500, PI)
NASA Earth System Science Fellowship, Climatic water-balance and the
dynamics of piñon-juniper woodlands, UNM, 2000 ($14,000, PI)
Sevilleta LTER Research Fellowship, 1998, 2000 ($2,000 each summer)
Graduate Research Allocations Committee, University of New Mexico, Fall 1999
($250)
Awards and Fellowships
NSF Biocomplexity Fellowship, UNM, 2001-2002
NSF Graduate Research Training Fellowship in Ecological Complexity, UNM,
1997-2000
Best Biology Teaching Assistant Award, UNM, Fall 1998
Peer-reviewed Publications
1. Kerkhoff, A.J. and B.J. Enquist. 2008. Multiplicative by nature: why
logarithmic transformation is necessary in allometry. Journal of Theoretical
Biology. In Press.
2. Bryant, J., C. Lamanna, H. Morlon, A.J. Kerkhoff, B.J. Enquist, and J.L.
Green. 2008. Microbes on mountainsides: Contrasting elevational patterns
of bacterial and plant diversity. Proceedings of the National Academy of
Sciences of the USA 105-suppl. 1: 11505-11511.
3. Enquist, B.J., A.J. Kerkhoff, S.C. Stark, N.G. Swenson, M.C. McCarthy,
and C.A. Price. 2007. A general integrative model for scaling plant growth
and functional trait spectra. Nature 449:218-222.
4. Decker, E.H., M.E. Moses, and A.J. Kerkhoff. 2007. Global patterns of city
size distributions and their fundamental drivers. PLoS-One 2:e934.
5. Kerkhoff, A.J., and B.J. Enquist. 2007. Implications of scaling approaches
for understanding resilience and reorganization in ecosystems.
BioScience 57:489-499.
6. White, E.P., S.K.M. Ernest, A.J. Kerkhoff, and B.J. Enquist. 2007. The
relationship between size and abundance in ecology. Trends in Ecology
and Evolution 22:323-330.
7. McCarthy, M.C., B.J. Enquist, and A.J. Kerkhoff. 2007. Organ partitioning
and allocation across the seed plants: Assessing the relative importance
of phylogeny and function. International Journal of Plant Sciences
168:751-761.
8. Weiser, M.D., B.J. Enquist, B. Boyle, T.J. Killeen, P.M. Jorgensen, G.
Fonseca, M. Jennings, A.J. Kerkhoff, T. Lacher, A. Monteagudo, M.P.
Nunez Vargas, O.L. Phillips, N.G. Swenson, and R. Vasquez Martinez.
2007. Latitudinal patterns of range size and species richness in New
World woody plants. Global Ecology and Biogeography 16:679-688.
9. Enquist, B.J., A.P. Allen, J.H. Brown, J.F. Gillooly, A.J. Kerkhoff, K.J.
Niklas, C.A. Price, and G.B. West. 2007. Metabolic scaling in seedlings:Does the exception prove the rule? Nature 445:E9-E10.
10. Ballantyne, F. and A.J. Kerkhoff. 2007. The observed range for mean-
variance scaling exponents can be explained by reproductive correlation.
Oikos 116:174-180.
11. Enquist, B.J., A.J. Kerkhoff, T.E. Huxman, and E.P. Economo. 2007.
Adaptive differences in plant physiology and ecosystem invariants:
Insights from a metabolic scaling model. Global Change Biology 13:591-
609.
12. Kerkhoff, A.J., W.F. Fagan, J.J. Elser, and B.J. Enquist. 2006.
Phylogenetic and functional variation in the scaling of nitrogen and
phosphorus in the seed plants. American Naturalist 168:E103-E122.
13. Kerkhoff, A.J. and B.J. Enquist. 2006. Ecosystem allometry: the scaling of
nutrient stocks and primary productivity across plant communities.
Ecology Letters 9:419-427.
14. Kerkhoff, A.J., B.J. Enquist, W.F. Fagan, and J.J. Elser. 2005. Plant
allometry, ecological stoichiometry and the temperature-dependence of
terrestrial primary productivity. Global Ecology and Biogeography 14:585-
598.
15. Ballantyne, F. and A.J. Kerkhoff. 2005. Synchronous reproduction and
mean-variance scaling in model forest stands. Journal of Theoretical
Biology 235:373-380.
16. Economo, E.P., A.J. Kerkhoff, and B.J. Enquist. 2005. Allometric growth,
life history invariants and population energetics. Ecology Letters 8:353-
360.
17. Kay, A.D., Ashton, I., Gorokhova, E., Kerkhoff, A.J., Leiss, A., and
Litchman, E. 2005. Toward a stoichiometric framework for evolutionary
biology. Oikos 109:6-17.
18. Kerkhoff, A.J. Expectation, explanation, and masting. 2004. Evoutionary
Ecology Research 6:1003-1020.
19. Kerkhoff, A.J., S.N. Martens, and B.T. Milne. 2004. An ecological test of
Eagleson’s optimality hypotheses. Functional Ecology 18:404-413.
20. Kerkhoff, A.J., S.N. Martens, G.A. Shore, and B.T. Milne. 2004.
Contingent effects of water balance variation on tree cover density in
semiarid woodlands. Global Ecology and Biogeography 13:237-246.
21. Kerkhoff, A.J. and F. Ballantyne. 2003. The scaling of reproductive
variability in trees. Ecology Letters 6:850-856.
22. Kerkhoff, A.J., B.T. Milne, and D.S. Maehr. 2000. Toward a panther-
centered view of the forests of South Florida. Conservation Ecology 4(1):1
[online] URL:http://www.consecol.org/vol4/iss1/art1.
Manuscripts in Review
Price, C.A., A.J. Kerkhoff, and B.J. Enquist. Growth form, tissue density and the
allometry of plant water and dry matter content. In Review. New Phytologist.Manuscripts in Preparation
Kerkhoff, A.J., R.B. Bash, S. Minium, P.R. Shanbhag, and H. Itagaki. The
ontogenetic scaling of growth, metabolism, and nutrient assimilation in larval
tobacco hawkmoths, Manduca sexta. For Functional Ecology.
Herbert E., M.K. Moe, E.C. Thompson, A.G. Lodge, and A.J. Kerkhoff.
Phylogenetic and biogeographic patterns of host tree association in
Lepidopteran herbivore communities. For American Naturalist.
Kerkhoff, A.J., B.J. Enquist, and C. Lamanna. Metabolic allometries and N:P
stoichiometry of montane plant communities. For Global Change Biology.
Elser, J.J., B.J. Enquist, W.F. Fagan and A.J. Kerkhoff. Tansley Review: Plant
stoichiometry, scaling, and global change. For New Phytologist.
Commentaries and Reviews of Publications
Webb, C.T. 2007. What is the role of ecology in understanding ecosystem
resilience? BioScience 57:470-471.
Hedin, L. 2004. Global organization of terrestrial plant nutrient interactions.
Proceedings of the National Academy of Sciences of the U.S.A. 101:10849-
10850.
Westoby, M. and I.J. Wright. 2006. Land-plant ecology on the basis of functional
traits. Trends in Ecology and Evolution 21:261-168.
Professional Service
Reviewer, 1999-present
American Naturalist, Biological Reviews, BioScience, Conservation Ecology,
Ecology, Ecology Letters, Ecoscience, Ecological Modelling, Evolution,
Functional Ecology, Global Ecology and Biogeography, Global Change
Biology, Journal of Ecology, Journal of Theoretical Biology, Landscape
Ecology, New Phytologist, Oikos, Proceedings of the Royal Society of
London B, Tree Physiology, Trends in Ecology and Evolution
Seminars Coordinator, 2006-2008, Kenyon College Biology Department
Biology Journal Club Advisor, 2006-2008, Kenyon College
Faculty Lectureships Committee, 2006-present, Kenyon College
Student-Faculty Senate, 2007-2008, Kenyon College
Committee Member, 2003, International Biogeography Society, Development
and Finance Committee
Committee Chair and Reviewer, 1998-2001, UNM Biology Graduate Resource
Allocations Committee
Graduate Student Representative, 2000-2001, UNM Biology Undergraduate
Policy Committee
Memberships
American Association for the Advancement of Science, Ecological Society of
America, American Society of Naturalists, International Biogeography Society,
American Institute of Biological SciencesInvited Presentations
March 2008. Department of Botany, Miami University, Oxford, OH.
October 2007. Department of Biology, West Virginia University, Morgantown,
WV.
November 2006. Kellogg Biological Station, Michigan State University, Hickory
Corners, MI.
May 2006. Department of Evolution, Ecology and Organismal Biology, Ohio State
University, Columbus, OH.
November 2004. Department of Biology, Kenyon College, Gambier, OH.
April 2004. Department of Biology, University of Texas, El Paso.
April 2004. Biocomplexity Seminar. University of New Mexico.
October 2003. Resilience and change in ecosystems workshop. The Santa Fe
Institute, Santa Fe, NM.
November 2003. Department of Ecology and Evolutionary Biology, University of
Arizona, Tucson, AZ.
June 2003. Rocky Mountain Biological Laboratory. Gothic, CO.
Contributed Presentations and Abstracts
Kerkhoff, A.J. and B.J. Enquist. August 2006. Metabolic allometries and the N:P
stoichiometry of montane plant communities. Poster presentation. Ecological
Society of America annual meeting. Memphis, TN.
Kerkhoff, A.J. and B.J. Enquist. July 2006. Metabolic allometries and the N:P
stoichiometry of montane plant communities. Poster presentation. Gordon
Research Conference, The Metabolic Basis of Ecology and Evolution.
Lewiston, ME.
Kerkhoff, A.J., W.F. Fagan, J.J. Elser, and B.J. Enquist. August 2005. N:P
stoichiometry of land plants: integration and coordination of the composition
of leaves, stems, roots and reproductive structures. Oral presentation.
Ecological Society of America annual meeting. Montreal, QC.
Enquist, B.J., A.J. Kerkhoff, B.L. Boyle, M.C. McCarthy, and N.G. Swenson.
August 2005. Assessing local and global constraints on plant community
structure: Quantifying small-scale distributions of size, functional traits, and
phylogenetic affiliations. Poster presentation. Ecological Society of America
annual meeting. Montreal, QC.
Pither, J., A.J. Kerkhoff, and B.J. Enquist. August 2005. Tolerance-differentiation
among species: a realistic and parsimonious resolution to the niche versus
neutrality debate? Oral presentation. Ecological Society of America annual
meeting. Montreal, QC.
Kerkhoff, A.J. and B.J. Enquist. February 2005. Ecosystem allometry: the scaling
of structure and function in whole plant communities. Poster presentation.
Graduate student recruitment week. University of Arizona Department of
Ecology and Evolutionary Biology.
Kerkhoff, A.J. July 2004. Ecosystem allometry: the scaling of structure and
function in whole plant communities. Poster presentation. Woodstoich 2004,
Finse, Norway.Kerkhoff, A.J. and B.J. Enquist. August 2004. Ecosystem allometry: the scaling of structure and function in whole plant communities. Oral presentation. Ecological Society of America annual meeting. Portland, OR. Enquist, B.J. and A.J. Kerkhoff. August 2004. From plant functional traits to communities - insights from a general energetic and stoichiometric scaling model. Oral presentation. Ecological Society of America annual meeting. Portland, OR. Kerkhoff, A.J. July 2004. Ecosystem allometry: the scaling of structure and function in whole plant communities. Poster presentation. Gordon Research Conference: The Metabolic Basis of Ecology, Lewiston, ME. Kerkhoff, A.J. August 2003. Plant allometry, ecological stoichiometry, and the scaling of terrestrial primary production. Oral presentation. Southwest Association of Biologists annual meeting. Portal, AZ. Kerkhoff, A.J., B.J. Enquist, W.F. Fagan, and J.J. Elser. August 2003. Latitudinal patterns in plant stoichiometry and the scaling of primary production. Oral presentation. Ecological Society of America annual meeting. Savannah, GA. Ballantyne, F. and A.J. Kerkhoff. August 2003. Masting, variance, and the mean. Oral presentation. Ecological Society of America annual meeting. Savannah, GA. Economo, E.P., A.J. Kerkhoff, and B.J. Enquist. August 2003. Allometric life histories and population energetics. Oral presentation. Ecological Society of America annual meeting. Savannah, GA. Kerkhoff, A.J., B.J. Enquist, W.F. Fagan, and J.J. Elser. January 2003. Latitudinal patterns of productivity and plant stoichiometry. Poster presentation. International Biogeography Society meeting. Mesquite NV. Weiser, M.D., B.J. Enquist, B. Boyle, E.P. Economo, A.J. Kerkhoff, S. Reddy, and C.O. Webb. January 2003. Species range size distributions of New World plant assemblages. Poster presentation. International Biogeography Society meeting. Mesquite, NV. Kerkhoff, A.J., and F. Ballantyne. October 2002. Invariant scaling of reproductive variability in trees. Oral presentation. Southwest Assoiciation of Biologists meeting. Portal, AZ. Kerkhoff, A.J. August 2002. Masting by chance or necessity: the reproductive variability of perennial plant populations. Oral presentation. Ecological Society of America annual meeting. Tucson, AZ. Kerkhoff, A.J., S.N. Martens, and B.T. Milne. August 2000. Ecological optimality in semi-arid woodlands: testing Eagleson’s ecohydrological equilibrium. Oral presentation, Ecological Society of America annual meeting. Snowbird, UT. Kerkhoff, A.J., S.N. Martens, and B.T. Milne. April 2000. Ecological optimality in a landscape. Poster presentation, International Association for Landscape Ecology, U.S., annual meeting. Ft. Lauderdale, FL. Milne, B.T., C. Restrepo, and A.J. Kerkhoff. August 1999. Multiscale assessment of structure and noise in species range maps. Invited oral presentation, Ecological Society of America annual meeting. Spokane, WA.
Kerkhoff, A.J., B.T. Milne, and D.S. Maehr. July 1999. Florida panthers: random walkers, or Levy flyers? Oral presentation, International Association for Landscape Ecology world congress. Snowmass, CO. Restrepo, C., A.J. Kerkhoff, and B.T. Milne. July 1999. Landslides and the large- scale organization of tropical montane ecosystems. Poster presentation, International Association for Landscape Ecology world congress, Snowmass, CO. Kerkhoff, A.J., and B.T. Milne. August 1997. Toward a panther-centered view of the forests of south Florida. Poster presentation, Ecological Society of America annual meeting. Albuquerque, NM. Decker, E.H., B.T. Milne, A.J. Kerkhoff, and P.T. Hraber. March 1997. Direct probability mapping by stochastically induced percolation. Poster presentation, International Association for Landscape Ecology, U.S., annual meeting. Durham, NC Kerkhoff, A.J., B.T. Milne, and D.S. Maehr. March 1997. Developing a panther- centered view of the south Florida landscape. Oral Presentation, International Association for Landscape Ecology, U.S., annual meeting. Durham, NC.
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