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The Handbook: Prevention and Control of Wildlife
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9-1-1994

Coyotes
Jeffrey S. Green
Assistant Regional Director USDA-APHIS Animal Damage Control

Green, Jeffrey S., "Coyotes" (1994). The Handbook: Prevention and Control of Wildlife Damage. Paper 34.
http://digitalcommons.unl.edu/icwdmhandbook/34

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Jeffrey S. Green
Assistant Regional Director
USDA-APHIS-
                                            COYOTES
Animal Damage Control
Lakewood, Colorado 80228

F. Robert Henderson
Extension Specialist
Animal Damage Control
Kansas State University
Manhattan, Kansas 66506-1600

Mark D. Collinge
State Director
USDA-APHIS-
Animal Damage Control
Boise, Idaho 83705

Fig. 1. Coyote, Canis latrans

Damage Prevention and                       Shed lambing, kidding, and calving            Toxicants
                                              usually reduce coyote predation.
Control Methods                                                                           M-44 ejector devices for use with
                                            Remove carrion to help limit coyote             sodium cyanide-loaded plastic
Exclusion                                     populations.                                  capsules. They are most effective
Produce livestock in confinement.           Frightening Agents and                          during cold weather (fall to spring).
                                            Repellents                                    Livestock protection collars (LPC)
Herd livestock into pens at night.
                                            Guarding dogs: Some dogs have                    containing Compound 1080
Exclusion fences (net-wire and/or                                                            (sodium monofluoroacetate) are
  electric), properly constructed and         significantly reduced coyote
                                              predation.                                     registered for use only in certain
  maintained, can aid significantly in                                                       states.
  reducing predation.                       Donkeys and llamas: Some are
                                              aggressive toward canines and have          Fumigants
Cultural Methods and
Habitat Modification                          reduced coyote predation.                   Gas cartridges are registered as a
                                            Sonic and visual repellents: Strobe             burrow (den) fumigant.
Select pastures that have a lower
   incidence of predation to reduce           lights, sirens, propane cannons, and        Trapping
   exposure of livestock to predation.        others have reduced predation on
                                              both sheep and calves.                      Leghold traps (Nos. 3 and 4) are
Herding of livestock generally reduces                                                      effective and are the most versatile
  predation due to human presence           Chemical odor and taste repellents:
                                                                                            control tool.
  during the herding period.                  None have shown sufficient
                                              effectiveness to be registered for          Snares are effective where coyotes pass
Change lambing, kidding, and calving          use.                                          through or under net-wire fences
  seasons.                                                                                  and in trail sets.

                                         PREVENTION AND CONTROL OF WILDLIFE DAMAGE — 1994
                                         Cooperative Extension Division
                                         Institute of Agriculture and Natural Resources
                                         University of Nebraska - Lincoln
                                         United States Department of Agriculture
                                         Animal and Plant Health Inspection Service
                                         Animal Damage Control
                                                                                                                                    C-51
                                         Great Plains Agricultural Council
                                         Wildlife Committee
Shooting                                    Aerial hunting is effective in removing     Other Methods
                                                     coyotes where terrain, ground
       Shooting from the ground is effective.
                                                     cover, vegetation, regulations, and       Denning: Remove adult coyotes and/
       Use rabbit distress calls or mimic            landownership conditions permit.            or their young from dens.
         howling or other coyote sounds to
                                                   Hunting with dogs is effective for trail-
         bring coyotes within shooting
                                                     ing coyotes from kill sites, locating
         distance.
                                                     dens, running coyotes, and assisting
                                                     with aerial hunting or calling.

       Identification                              Habitat                                     weaker lambs are also present. Usu-
                                                                                               ally, the stronger lamb is on the
       In body form and size, the coyote           Many references indicate that coyotes       periphery and is more active, making
       (Canis latrans) resembles a small collie    were originally found in relatively         it more prone to attack than a weaker
       dog, with erect pointed ears, slender       open habitats, particularly the grass-      lamb that is at the center of the flock
       muzzle, and a bushy tail (Fig. 1).          lands and sparsely wooded areas of          and relatively immobile.
       Coyotes are predominantly brownish          the western United States. Whether or       Coyote predation on livestock is gener-
       gray in color with a light gray to          not this was true, coyotes have             ally more severe during early spring
       cream-colored belly. Color varies           adapted to and now exist in virtually       and summer than in winter for two
       greatly, however, from nearly black to      every type of habitat, arctic to tropic,    reasons. First, sheep and cows are usu-
       red or nearly white in some individu-       in North America. Coyotes live in           ally under more intensive manage-
       als and local populations. Most have        deserts, swamps, tundra, grasslands,        ment during winter, either in feedlots
       dark or black guard hairs over their        brush, dense forests, from below sea        or in pastures that are close to human
       back and tail. In western states, typical   level to high mountain ranges, and at       activity, thus reducing the opportunity
       adult males weigh from 25 to 45             all intermediate altitudes. High densi-     for coyotes to take livestock. Second,
       pounds (11 to 16 kg) and females from       ties of coyotes also appear in the sub-     predators bear young in the spring and
       22 to 35 pounds (10 to 14 kg). In the       urbs of Los Angeles, Pasadena,              raise them through the summer, a pro-
       East, many coyotes are larger than          Phoenix, and other western cities.          cess that demands increased nutri-
       their western counterparts, with males                                                  tional input, for both the whelping and
       averaging about 45 pounds (14 kg) and                                                   nursing mother and the growing
       females about 30 pounds (13 kg).            Food Habits
                                                                                               young. This increased demand corre-
       Coyote-dog and coyote-wolf hybrids                                                      sponds to the time when young sheep
                                                   Coyotes often include many items in
       exist in some areas and may vary                                                        or beef calves are on pastures or range-
                                                   their diet. Rabbits top the list of their
       greatly from typical coyotes in size,                                                   land and are most vulnerable to attack.
                                                   dietary components. Carrion, rodents,
       color, and appearance. Also, coyotes in                                                 Coyote predation also may increase
                                                   ungulates (usually fawns), insects
       the New England states may differ in                                                    during fall when young coyotes dis-
                                                   (such as grasshoppers), as well as live-
       color from typical western coyotes.                                                     perse from their home ranges and
                                                   stock and poultry, are also consumed.
       Many are black, and some are reddish.                                                   establish new territories.
                                                   Coyotes readily eat fruits such as
       These colorations may partially be due      watermelons, berries, and other veg-
       to past hybridization with dogs and         etative matter when they are available.     General Biology,
       wolves. True wolves are also present        In some areas coyotes feed on human         Reproduction, and
       in some areas of coyote range, particu-     refuse at dump sites and take pets
       larly in Canada, Alaska, Montana,
                                                                                               Behavior
                                                   (cats and small dogs).
       northern Minnesota, Wisconsin, and
                                                   Coyotes are opportunistic and gener-        Coyotes are most active at night and
       Michigan. Relatively few wolves
                                                   ally take prey that is the easiest to       during early morning hours (especially
       remain in the southern United States
                                                   secure. Among larger wild animals,          where human activity occurs), and
       and Mexico.
                                                   coyotes tend to kill young, inexperi-       during hot summer weather. Where
                                                   enced animals, as well as old, sick, or     there is minimal human interference
       Range                                                                                   and during cool weather, they may be
                                                   weakened individuals. With domestic
                                                   animals, coyotes are capable of catch-      active throughout the day.
       Historically, coyotes were most com-
                                                   ing and killing healthy, young, and in      Coyotes bed in sheltered areas but do
       mon on the Great Plains of North
                                                   some instances, adult prey. Prey selec-     not generally use dens except when
       America. They have since extended
                                                   tion is based on opportunity and a          raising young. They may seek shelter
       their range from Central America to
                                                   myriad of behavioral cues. Strong,          underground during severe weather
       the Arctic, including all of the United
                                                   healthy lambs are often taken from a        or when closely pursued. Their physi-
       States (except Hawaii), Canada, and
                                                   flock by a coyote even though smaller,      cal abilities include good eyesight and
       Mexico.

C-52
hearing and a keen sense of smell.           feeding and caring for the young. Coy-      the site. The quantity of sheep or calf
Documented recoveries from severe            otes commonly hunt as singles or            remains left after a kill vary widely
injuries are indicative of coyotes’          pairs; extensive travel is common in        depending on how recently the kill
physical endurance. Although not as          their hunting forays. They will hunt in     was made, the size of the animal
fleet as greyhound dogs, coyotes have        the same area regularly, however, if        killed, the weather, and the number
been measured at speeds of up to 40          food is plentiful. They occasionally        and species of predators that fed on
miles per hour (64 km/hr) and can            bury food remains for later use.            the animal.
sustain slower speeds for several miles
                                             Pups begin emerging from their den          One key in determining whether a
(km).
                                             by 3 weeks of age, and within 2             sheep or calf was killed by a predator
Distemper, hepatitis, parvo virus, and       months they follow adults to large          is the presence or absence of subcuta-
mange (caused by parasitic mites) are        prey or carrion. Pups normally are          neous (just under the skin) hemor-
among the most common coyote dis-            weaned by 6 weeks of age and fre-           rhage at the point of attack. Bites to a
eases. Rabies and tularemia also occur       quently are moved to larger quarters        dead animal will not produce hemor-
and may be transmitted to other ani-         such as dense brush patches and/or          rhage, but bites to a live animal will. If
mals and humans. Coyotes harbor              sinkholes along water courses. The          enough of the sheep carcass remains,
numerous parasites including mites,          adults and pups usually remain              carefully skin out the neck and head to
ticks, fleas, worms, and flukes. Mortal-     together until late summer or fall          observe tooth punctures and hemor-
ity is highest during the first year of      when pups become independent.               rhage around the punctures. Talon
life, and few survive for more than 10       Occasionally pups are found in groups       punctures from large birds of prey will
to 12 years in the wild. Human activity      until the breeding season begins.           also cause hemorrhage, but the loca-
is often the greatest single cause of                                                    tion of these is usually at the top of the
                                             Coyotes are successful at surviving
coyote mortality.                                                                        head, neck, or back. This procedure
                                             and even flourishing in the presence of
                                                                                         becomes less indicative of predation as
Coyotes usually breed in February and        people because of their adaptable
                                                                                         the age of the carcass increases or if the
March, producing litters about 9             behavior and social system. They typi-
                                                                                         remains are scanty or scattered.
weeks (60 to 63 days) later in April and     cally display increased reproduction
May. Females sometimes breed during          and immigration in response to              Coyotes, foxes, mountain lions, and
the winter following their birth, par-       human-induced population reduction.         bobcats usually feed on a carcass at the
ticularly if food is plentiful. Average                                                  flanks or behind the ribs and first con-
litter size is 5 to 7 pups, although up to                                               sume the liver, heart, lungs, and other
13 in a litter has been reported. More       Damage and Damage                           viscera. Mountain lions often cover a
than one litter may be found in a single     Identification                              carcass with debris after feeding on it.
den; at times these may be from                                                          Bears generally prefer meat to viscera
females mated to a single male. As           Coyotes can cause damage to a variety       and often eat first the udder from lac-
noted earlier, coyotes are capable of        of resources, including livestock, poul-    tating ewes. Eagles skin out carcasses
hybridizing with dogs and wolves, but        try, and crops such as watermelons.         on larger animals and leave much of
reproductive dysynchrony and                 They sometimes prey on pets and are a       the skeleton intact. With smaller ani-
behaviors generally make it unlikely.        threat to public health and safety when     mals such as lambs, eagles may bite off
Hybrids are fertile, although their          they frequent airport runways and           and swallow the ribs. Feathers and
breeding seasons do not usually corre-       residential areas, and act as carriers of   “whitewash” (droppings) are usually
spond to those of coyotes.                   rabies. Usually, the primary concern        present where an eagle has fed.
                                             regarding coyotes is predation on live-
Coyote dens are found in steep banks,                                                    Coyotes may kill more than one ani-
                                             stock, mainly sheep and lambs. Preda-
rock crevices, sinkholes, and under-                                                     mal in a single episode, but often will
                                             tion will be the focus of the following
brush, as well as in open areas. Usu-                                                    only feed on one of the animals.
                                             discussion.
ally their dens are in areas selected for                                                Coyotes typically attack sheep at the
protective concealment. Den sites are        Since coyotes frequently scavenge on        throat, but young or inexperienced
typically located less than a mile (km)      livestock carcasses, the mere presence      coyotes may attack any part of the
from water, but may occasionally be          of coyote tracks or droppings near a        body. Coyotes usually kill calves by
much farther away. Coyotes will often        carcass is not sufficient evidence that     eating into the anus or abdominal area.
dig out and enlarge holes dug by             predation has taken place. Other evi-
                                                                                         Dogs generally do not kill sheep or
smaller burrowing animals. Dens vary         dence around the site and on the car-
                                                                                         calves for food and are relatively
from a few feet (1 m) to 50 feet (15 m)      cass must be carefully examined to aid
                                                                                         indiscriminate in how and where they
and may have several openings.               in determining the cause of death.
                                                                                         attack. Sometimes, however, it is
                                             Signs of a struggle may be evident.
Both adult male and female coyotes                                                       difficult to differentiate between dog
                                             These may include scrapes or drag
hunt and bring food to their young for                                                   and coyote kills without also looking
                                             marks on the ground, broken vegeta-
several weeks. Other adults associated                                                   at other sign, such as size of tracks
                                             tion, or blood in various places around
with the denning pair may also help in                                                   (Fig. 2) and spacing and size of canine

                                                                                                                                      C-53
tooth punctures. Coyote tracks tend to
       be more oval-shaped and compact
       than those of common dogs. Nail
       marks are less prominent and the
       tracks tend to follow a straight line
       more closely than those of dogs. The
       average coyote’s stride at a trot is 16 to
       18 inches (41 to 46 cm), which is typi-
       cally longer than that of a dog of simi-
       lar size and weight. Generally, dogs                                                                                        3"
       attack and rip the flanks, hind quar-
       ters, and head, and may chew ears.
       The sheep are sometimes still alive but                                                  5"               Coyote
       may be severely wounded.
       Accurately determining whether or
       not predation occurred and, if so, by
       what species, requires a considerable
       amount of knowledge and experience.
       Evidence must be gathered, pieced                                  Wolf
       together, and then evaluated in light of
       the predators that are in the area, the
       time of day, the season of the year, and
       numerous other factors. Sometimes
       even experts are unable to confirm the
       cause of death, and it may be neces-
       sary to rely on circumstantial informa-
       tion. For more information on this
       subject, refer to the section Procedures
       for Evaluating Predation on Livestock                                      2 1/2"
                                                                                                                                    4"
       and Wildlife, in this book.
                                                                   Red fox
       Legal Status                                                                                            Large dog

       The status of coyotes varies depending       Fig. 2. Footprints of canid predators
       on state and local laws. In some states,
       including most western states, coyotes
       are classified as predators and can be       Federal statutes that pertain to wildlife        Damage Prevention and
       taken throughout the year whether or         damage control include the Federal
       not they are causing damage to live-         Insecticide, Fungicide, and Rodenti-
                                                                                                     Control Methods
       stock. In other states, coyotes may be       cide Act (FIFRA), which deals with
                                                                                                     For managing coyote damage, a vari-
       taken only during specific seasons and       using toxicants, and the Airborne
                                                                                                     ety of control methods must be avail-
       often only by specific methods, such as      Hunting Act, which regulates aerial
                                                                                                     able since no single method is effective
       trapping. Night shooting with a spot-        hunting.
                                                                                                     in every situation. Success usually
       light is usually illegal. Some state laws
                                                    Laws regulating coyote control are not           involves an integrated approach, com-
       allow only state or federal agents to
                                                    necessarily uniform among states or              bining good husbandry practices with
       use certain methods (such as snares) to
                                                    even among counties within a state,              effective control methods for short
       take coyotes. Some states have a provi-
                                                    and they may change frequently. A                periods of time. Regardless of the
       sion for allowing the taking of pro-
                                                    1989 Supreme Court action established            means used to stop damage, the focus
       tected coyotes (usually by special
                                                    that it was not legal to circumvent the          should be on damage prevention and
       permit) when it has been documented
                                                    laws relative to killing predators, even         control rather than elimination of coy-
       that they are preying on livestock. In
                                                    to protect personal property (livestock)         otes. It is neither wise nor practical to
       some instances producers can apply
                                                    from predation.                                  kill all coyotes. It is important to try to
       control methods, and in others, control
                                                                                                     prevent coyotes from killing calves or
       must be managed by a federal or state
                                                                                                     sheep for the first time. Once a coyote
       agent. Some eastern states consider the
                                                                                                     has killed livestock, it will probably
       coyote a game animal, a furbearer, or a
                                                                                                     continue to do so if given the
       protected species.

C-54
opportunity. Equally important is
taking action as quickly as possible to
stop coyotes from killing after they
start.
                                                                                                    1/4" x 4 1/2" carriage bolt
Exclusion                                                                                           galvanized woven wire

Most coyotes readily cross over,
under, or through conventional live-
stock fences. A coyote’s response to a
fence is influenced by various factors,
                                                                        72" galvanized                 3" treated pole
including the coyote’s experience and                                                                       7' long
                                                                             fencing
motivation for crossing the fence. Total                                 6" stays
                                                          69"
exclusion of all coyotes by fencing,                                     4" vertical
especially from large areas, is highly                                       spacing at top
                                                                        1 1/2" at the
unlikely since some eventually learn to                           56"        bottom
either dig deeper or climb higher to
defeat a fence. Good fences, however,
can be important in reducing preda-
tion, as well as increasing the effective-
ness of other damage control methods
(such as snares, traps, or guarding                                          23" apron
animals).                                                                    (old fencing)

Recent developments in fencing equip-
ment and design have made this
technique an effective and economi-
cally practical method for protecting                                                                 28"
sheep from predation under some
grazing conditions. Exclusion fencing
may be impractical in western range
sheep ranching operations.
                                             Fig. 3. Barrier fence with wire overhang and
Net-Wire Fencing. Net fences in              buried apron.
good repair will deter many coyotes
from entering a pasture. Horizontal
spacing of the mesh should be less
than 6 inches (15 cm), and vertical          Electric Fencing. Electric fencing,              The number of spacings between wires
spacing less than 4 inches (10 cm). Dig-     used for years to manage livestock, has          varies considerably. A fence of 13
ging under a fence can be discouraged        recently been revolutionized by the              strands gave complete protection to
by placing a barbed wire at ground           introduction of new energizers and               sheep from coyote predation in tests at
level or using a buried wire apron           new fence designs from Australia and             the USDA’s US Sheep Experiment Sta-
(often an expensive option). The fence       New Zealand. The chargers, now also              tion (Fig. 4). Other designs of fewer
should be about 5 1/2 feet (1.6 m) high      manufactured in the United States,               wires were effective in some studies,
to discourage coyotes from jumping           have high output with low impedance,             ineffective in others.
over it. Climbing can usually be pre-        are resistant to grounding, present a
                                                                                              The amount of labor and installation
vented by adding a charged wire at           minimal fire hazard, and are generally
                                                                                              techniques required vary with each
the top of the fence or installing a wire    safe for livestock and humans. The
                                                                                              type of fencing. High-tensile wire
overhang.                                    fences are usually constructed of
                                                                                              fences require adequate bracing at cor-
                                             smooth, high-tensile wire stretched to
Barrier fences with wire overhangs                                                            ners and over long spans. Electric fenc-
                                             a tension of 200 to 300 pounds (90 to
and buried wire aprons were tested in                                                         ing is easiest to install on flat, even
                                             135 kg). The original design of electric
Oregon and found effective in keeping                                                         terrain. Labor to install a high-tensile
                                             fences for controlling predation con-
coyotes out of sheep pastures (Fig. 3).                                                       electric fence may be 40% to 50% less
                                             sisted of multiple, alternately charged
The construction and materials for                                                            than for a conventional livestock fence.
                                             and grounded wires, with a charged
such fencing are usually expensive.          trip wire installed just above ground            Labor to keep electric fencing func-
Therefore, fences of this type are rarely    level about 8 inches (20 cm) outside the         tional can be significant. Tension of the
used except around corrals, feedlots,        main fence to discourage digging.                wires must be maintained, excessive
or areas of temporary sheep confine-         Many recent designs have every wire              vegetation under the fence must be
ment.                                        charged.                                         removed to prevent grounding, dam-

                                                                                                                                          C-55
age from livestock and wildlife must
                                                                                  Fiberglass
                                                                                  line post                     be repaired, and the charger must be
                                                                                                                checked regularly to ensure that it is
                                                                                                                operational.
                                                                                                                Coyotes and other predators occasion-
                                                                    66'                                         ally become “trapped” inside electric
                                                                                                                fences. These animals receive a shock
                                                             Fiberglass                                 6'
                                              22'            stays                                              as they enter the pasture and subse-
                                       11'                                                                      quently avoid approaching the fence to
                                                                                                                escape. In some instances the captured
                                                                                                                predator may be easy to spot and
                  Fiberglass                                                                                    remove from the pasture, but in
                  line post                                                                                     others, particularly in large pastures
                                                                                                                with rough terrain, the animal may be
       Charged wire                                                                                             difficult to remove.
                          + 8"
       Ground wire
                          - 8"                                                                                  Electric Modification of Existing
                          + 8"                                                                     Fiberglass   Fences. The cost to completely
                                                                                                   stay
                          - 8"                                                                                  replace old fences with new ones,
                          + 8"                                                                                  whether conventional or electric, can
                          -   8"                                      Trip wire                                 be substantial. In instances where
                          +   8"                                                                                existing fencing is in reasonably good
                          -                            8"
                              8"                                                                                condition, the addition of one to sev-
                          +   4"                               6"
                          -   4"                                            Note:                               eral charged wires can significantly
                          +   4"                                            Drawing not to scale
                                                                                                                enhance the predator-deterring ability
                          -   4"
                                                                                                                of the fence and its effectiveness for
                                                                                                                controlling livestock (Fig. 5). A
                                          Ground                                                                charged trip wire placed 6 to 8 inches
                                          level
                                                                                                                (15 to 230 cm) above the ground about
                                                                                                                8 to 10 inches (20 to 25 cm) outside the
       Fig. 4. High-tensile, electric, antipredator fence.                                                      fence is often effective in preventing
                                                                                                                coyotes from digging and crawling un-
                                                                                                                der. This single addition to an existing
                                                                                                                fence is often the most effective and
                                                                                                                economical way to fortify a fence
                                                                                                                against coyote passage.
                                                                                                                If coyotes are climbing or jumping a
                                                                                                                fence, charged wires can be added to
                                                                                                                the top and at various intervals. These
                                                                                                                wires should be offset outside the
                                                                                                                fence. Fencing companies offer offset
                                                                                                                brackets to make installation relatively
                                                                                                                simple. The number of additional
                                                                                                                wires depends on the design of the
                                                                                                                original fence and the predicted habits
                                                                                                                of the predators.
                                                                                                                Portable Electric Fencing. The
                                                                                     Outrigger post
                                                                                                                advent of safe, high-energy chargers
                                                                                     with four wires
                                                                                                                has led to the development of a variety
                                                                                                                of portable electric fences. Most are
                                                                                                                constructed with thin strands of wire
                                                                                                                running through polyethylene twine or
       Fig. 5. Existing woven-wire livestock fence modified with electrified wire.
                                                                                                                ribbon, commonly called polywire or
                                                                                                                polytape. The polywire is available in
                                                                                                                single and multiple wire rolls or as
                                                                                                                mesh fencing of various heights. It can
                                                                                                                be quickly and easily installed to serve

C-56
as a temporary corral or to partition         As a final note, fences can pose prob-      Counting sheep and calves regularly is
off pastures for controlled grazing.          lems for wildlife. Barrier fences in par-   important in large pastures or areas
                                              ticular exclude not only predators, but     with heavy cover where dead livestock
Perhaps the biggest advantage of port-
                                              also many other wildlife species. This      could remain unnoticed. It is not
able electric fencing is the ability to set
                                              fact should be considered where fenc-       unusual for producers who do not
up temporary pens to hold livestock at
                                              ing intersects migration corridors for      regularly count their sheep to suffer
night or during other predator control
                                              wildlife. Ungulates such as deer may        fairly substantial losses before they
activities. Portable fencing increases
                                              attempt to jump fences, and they occa-      realize there is a problem. Determining
livestock management options to
                                              sionally become entangled in the top        with certainty whether losses were due
avoid places or periods of high preda-
                                              wires.                                      to coyotes or to other causes may
tion risk. Range sheep that are not
                                                                                          become impossible.
accustomed to being fenced, however,          Cultural Methods and Habitat
may be difficult to contain in a port-                                                    Season and Location of Lambing
                                              Modification
able fence.                                                                               or Calving. Both season and location
                                              At the present time, there are no docu-     of lambing and calving can signifi-
Fencing and Predation Manage-                 mented differences in the vulnerability     cantly affect the severity of coyote pre-
ment. The success of various types of
                                              of various breeds of sheep to coyote or     dation on sheep or calves. The highest
fencing in keeping out predators has          dog predation because there has been        predation losses of sheep and calves
ranged from poor to excellent. Density        very little research in this area. Gener-   typically occur from late spring
and behavior of coyotes, terrain and          ally, breeds with stronger flocking         through September due to the food
vegetative conditions, availability of        behaviors are less vulnerable to preda-     requirements of coyote pups. In the
prey, size of pastures, season of the         tors.                                       Midwest and East, some lambing or
year, design of the fence, quality of
                                              A possible cause of increased coyote        calving occurs between October and
construction, maintenance, and other
                                              predation to beef cattle calves is the      December, whereas in most of the
factors all interplay in determining
                                              increased use of cattle dogs in herding.    western states lambing or calving
how effective a fence will be. Fencing
                                              Cows herded by dogs may not be as           occurs between February and May. By
is most likely to be cost-effective where
                                              willing to defend newborn calves from       changing to a fall lambing or calving
the potential for predation is high,
                                              coyotes as those not accustomed to          program, some livestock producers
where there is potential for a high
                                              herding dogs.                               have not only been able to diversify
stocking rate, or where electric modifi-
                                                                                          their marketing program, but have
cation of existing fences can be used.        Flock or Herd Health. Healthy               also avoided having a large number of
Fencing can be effective when incorpo-        sheep flocks and cow/calf herds have        young animals on hand during periods
rated with other means of predation           higher reproductive rates and lower         when coyote predation losses are typi-
control. For example, combined use of         overall death losses. Coyotes often         cally highest.
guarding dogs and fencing has                 prey on smaller lambs. Poor nutrition
                                              means weaker or smaller young, with         Shortening lambing and calving peri-
achieved a greater degree of success
                                              a resultant increased potential for pre-    ods by using synchronized or group
than either method used alone. An
                                              dation. Ewes or cows in good condi-         breeding may reduce predation by
electric fence may help keep a guard-
                                              tion through proper nutrition will raise    producing a uniform lamb or calf crop,
ing dog in and coyotes out of a pas-
                                              stronger young that may be less vul-        thus reducing exposure of small live-
ture. If an occasional coyote does pass
                                              nerable to coyote predation.                stock to predation. Extra labor and
through a fence, the guarding dog can
                                                                                          facilities may be necessary, however,
keep it away from the livestock and           Record Keeping. Good record-                when birthing within a concentrated
alert the producer by barking.                keeping and animal identification sys-      period. Some producers practice early
Fencing can also be used to concen-           tems are invaluable in a livestock          weaning and do not allow young to go
trate predator activity at specific places    operation for several reasons. From the     to large pastures, thus reducing the
such as gateways, ravines, or other           standpoint of coyote predation,             chance of coyote losses. This also gives
areas where the animals try to gain           records help producers identify loss        orphaned and weak young a greater
access. Traps and snares can often be         patterns or trends to provide baseline      chance to survive.
set at strategic places along a fence to      data that will help determine what
                                              type and amount of coyote damage            The average beef cattle calf production
effectively capture predators. Smaller
                                              control is economically feasible.           is about 78% nationwide. First-calf
pastures are easier to keep free from
                                              Records also aid in identifying critical    heifers need human assistance to give
predators than larger ones encompass-
                                              problem areas that may require atten-       birth to a healthy calf about 40% of the
ing several square miles (km2).
                                              tion. They may show, for example, that      time. Cow/calf producers who aver-
Fencing is one of the most beneficial         losses to coyotes are high in a particu-    age 90% to 95% calf crops generally
investments in predator damage con-           lar pasture in early summer, thus high-     check their first-calf heifers every 2
trol and livestock management where           lighting the need for preventive            hours during calving. Also, most good
practical factors warrant its use.            control in that area.                       producers place first-calf heifers in

                                                                                                                                      C-57
small pastures (less than 160 acres [64     Some producers reason that coyotes              nerable to coyote predation than those
       ha]). When all cows are bred to pro-        are less likely to kill livestock if there is   in closer, more open, and smaller pas-
       duce calves in a short, discreet (e.g.      carrion available. This may be a valid          tures. In general, a relatively small,
       60-day) period, production typically        preventative measure if an adequate             open, tightly fenced pasture that can
       increases and predation losses              supply of carrion can be maintained             be kept under close surveillance is a
       decrease. The birth weight of calves        far away from livestock. If a coyote            good choice for birthing livestock that
       born to first-calf heifers can be de-       becomes habituated to a diet of live-           are likely targets of coyotes. Past expe-
       creased by using calving-ease bulls,        stock remains, however, it may turn to          rience with predators as well as
       thus reducing birthing complications        killing livestock in the absence of car-        weather and disease considerations
       that often lead to coyote predation.        casses. Wherever there is easily acces-         should also serve as guides in the
                                                   sible carrion, coyotes seem to gather           selection of birthing pastures.
       Producers who use lambing sheds or
                                                   and predation losses are higher. Con-
       pens for raising sheep and small pas-                                                       A factor not completely understood is
                                                   versely, where carrion is generally not
       tures or paddocks for raising cattle                                                        that, at times, coyotes and other preda-
                                                   available, losses are lower. A study in
       have lower predation losses than those                                                      tors will kill in one pasture and not in
                                                   Canada showed that the removal of
       who lamb or calve in large pastures or                                                      another. Therefore, changing pastures
                                                   livestock carcasses significantly
       on open range. The more human pres-                                                         during times of loss may reduce pre-
                                                   reduced overwinter coyote popula-
       ence around sheep, the lower the pre-                                                       dation. There may seem to be a rela-
                                                   tions and shifted coyote distributions
       dation losses. Confining sheep entirely                                                     tionship between size of pasture and
                                                   out of livestock areas.
       to buildings virtually eliminates preda-                                                    predator losses, with higher loss rates
       tion losses.                                Habitat Changes. Habitat features               reported in larger pastures. In reality,
                                                   change in some areas, depending on              loss rates may not be related as much
       Corrals. Although predation can
                                                   seasonal crop growth. Some cultivated           to pasture size as to other local condi-
       occur at any time, coyotes tend to kill
                                                   fields are devoid of coyotes during             tions such as slope, terrain, and human
       sheep at night. Confining sheep at
                                                   winter but provide cover during the             populations. Hilly or rugged areas are
       night is one of the most effective
                                                   growing season, and a corresponding             typically sparsely populated by hu-
       means of reducing losses to predation.
                                                   increase in predation on nearby live-           mans and are characterized by large
       Nevertheless, some coyotes and many
                                                   stock may occur.                                pastures. These conditions are ideal for
       dogs are bold enough to enter corrals
                                                   The creation of nearly 40 million acres         coyotes.
       and kill sheep. A “coyote-proof” corral
       is a wise investment. Coyotes are more      (16 million ha) of Conservation                 Sheep pastures that contain or are
       likely to attack sheep in unlighted cor-    Reserve Program (CRP) acres may                 adjacent to streams, creeks, and rivers
       rals than in corrals with lights. Even if   benefit many species of wildlife,               tend to have more coyote problems
       the corral fence is not coyote-proof, the   including predators. These acres har-           than pastures without such features.
       mere fact that the sheep are confined       bor prey for coyotes and foxes, and an          Water courses serve as hunting and
       reduces the risk of predation. Penning      increase in predator populations can            travel lanes for coyotes.
       sheep at night and turning them out at      reasonably be predicted. Clearing
                                                   away weeds and brush from CRP                   Herders. Using herders with sheep or
       mid-morning might reduce losses. In
                                                   areas may reduce predation problems             cattle in large pastures can help reduce
       addition, coyotes tend to be more
                                                   since predators usually use cover in            predation, but there has been a trend
       active and kill more sheep on foggy or
                                                   their approach to livestock. Generally,         away from herders in recent years
       rainy days than on sunny days. Keep-
                                                   the more open the area where live-              because of increasing costs and a
       ing the sheep penned on foggy or rainy
                                                   stock are kept, the less likely that            shortage of competent help. Neverthe-
       days may be helpful.
                                                   coyote losses will occur. Often junk            less, tended flocks or herds receive
       Aside from the benefits of livestock        piles are located near farmsteads.              closer attention than untended live-
       confinement, there are some problems        These serve as good habitat for rabbits         stock, particularly in large pastures,
       associated it. Costs of labor and mate-     and other prey and may bring coyotes            and problems can be solved before
       rials associated with building corrals,     into close proximity with livestock,            they become serious. We recommend
       herding livestock, and feeding live-        increasing the likelihood for opportu-          two herders per band of range sheep.
       stock must be considered. In addition,      nistic coyotes to prey on available live-       If herders aren’t used, daily or periodic
       the likelihood of increased parasite        stock. Removing junk piles may be a             checking of the livestock is a good hus-
       and disease problems may inhibit            good management practice.                       bandry practice.
       adoption of confinement as a method
       of reducing damage.                         Pasture Selection. If sheep or beef             Frightening Devices and
                                                   cattle are not lambed or calved in              Repellents
       Carrion Removal. Removal and                sheds or lots, the choice of birthing
       proper disposal of dead sheep and           pastures should be made with poten-             Frightening devices are useful for
       cattle are important since livestock car-   tial coyote predation problems in               reducing losses during short periods
       casses tend to attract coyotes,             mind. Lambs and calves in remote or             or until predators are removed. The
       habituating them to feed on livestock.      rugged pastures are usually more vul-           devices should not be used for long

C-58
periods of time when predation is not       Some producers use a radio tuned to
a problem. To avoid acclimation you         an all-night station to temporarily
can increase both the degree and dura-      deter coyotes, dogs, and other preda-
tion of effectiveness by varying the        tors.
position, appearance, duration, or fre-
                                            Vehicles. Parking cars or pickups in
quency of the frightening stimuli, or
                                            the area where losses are occurring of-
using them in various combinations.
                                            ten reduces predation temporarily.
Many frightening methods have been
                                            Effectiveness can be improved or
ridiculed in one way or another; never-
                                            extended by frequently moving the
theless, all of the techniques discussed
                                            vehicle to a new location. Some pro-
here have helped producers by saving
                                            ducers place a replica of a person in
livestock and/or buying some time to
                                            the vehicle when losses are occurring      Fig. 6. Electronic Guard frightening device
institute other controls.
                                            in the daylight. If predators continue
Lights. A study involving 100 Kansas        to kill with vehicles in place, the
                                                                                       Center developed a frightening device
sheep producers showed that using           vehicle serves as a comfortable blind in
                                                                                       called the Electronic Guard (EG) (Fig.
lights above corrals at night had the       which to wait and shoot offending
                                                                                       6). The EG consists of a strobe light
most marked effect on losses to coy-        predators.
                                                                                       and siren controlled by a variable
otes of all the devices examined. Out
                                            Propane Exploders. Propane                 interval timer that is activated at night
of 79 sheep killed by coyotes in corrals,
                                            exploders produce loud explosions at       with a photoelectric cell. In tests con-
only three were killed in corrals with
                                            timed intervals when a spark ignites a     ducted in fenced pastures, predation
lights. Nearly 40% of the producers in
                                            measured amount of propane gas. On         was reduced by about 89%. The device
the study used lights over corrals.
                                            most models, the time between explo-       is used in Kansas and other states to
There was some indication in the
                                            sions can vary from about 1 minute to      protect cows/calves from coyote pre-
study that sheep losses to dogs were
                                            15 minutes. Their effectiveness at         dation. Most research on the effective-
higher in lighted corrals, but the
                                            frightening coyotes is usually only        ness of this device, however, has been
sample size for dog losses was small
                                            temporary, but it can be increased by      done on sheep operations. Suggestions
and the results inconclusive. Most of
                                            moving exploders to different loca-        for using the unit differ for pastured
the producers (80%) used mercury
                                            tions and by varying the intervals be-     sheep and range operations.
vapor lights that automatically turned
                                            tween explosions. In general, the timer
on at dusk and off at dawn.                                                            To use the EG in fenced pastures (farm
                                            on the exploder should be set to fire
                                                                                       flocks):
Another advantage of lighted corrals is     every 8 to 10 minutes, and the location
that coyotes are more vulnerable when       should be changed every 3 or 4 days.       1. Place EGs above the ground on
they enter the lighted area. Coyotes        In cattle pastures, these devices should      fence posts, trees, or T-posts so they
often establish a fairly predictable pat-   be placed on rigid stands above the           can be heard and seen at greater
tern of killing. When this happens in a     livestock. Normally, the exploder             distances and to prevent livestock
lighted corral, it is possible for a pro-   should be turned on just before dark          from damaging them.
ducer to wait above or downwind of          and off at daybreak, unless coyotes are
                                                                                       2. Position EGs so that rain water can-
the corral and to shoot the coyote as it    killing livestock during daylight hours.
                                                                                          not enter them and cause a malfunc-
enters. Red or blue lights may make         Motion sensors are now available and
                                                                                          tion.
the ambush more successful since coy-       likely improve their effectiveness,
otes appear to be less frightened by        though it is still only temporary.         3. Locate EGs so that light can enter
them than by white lights.                  Exploders are best used to reduce             the photocell port or window. If
                                            losses until more permanent control or        positioned in deep shade, they may
Revolving or flashing the lights may
                                            preventive measures can be imple-             not turn on or off at the desired
enhance their effectiveness in frighten-
                                            mented. In about 24 coyote depreda-           times.
ing away predators. There is some
                                            tion complaints over a 2-year period in
speculation that the old oil lamps used                                                4. The number of EGs used to protect
                                            North Dakota, propane exploders
in highway construction repelled                                                          sheep in fenced pastures depends
                                            were judged to be successful in stop-
coyotes, presumably because of their                                                      on pasture size, terrain features, and
                                            ping or reducing predation losses until
flickering effect.                                                                        the amount and height of vegetation
                                            offending coyotes could be removed.
                                                                                          in or around the pasture. In general,
Bells and Radios. Some sheep pro-           “Success time” of the exploders ap-
                                                                                          at least two units should be used in
ducers place bells on some or all of        pears to depend a great deal on how
                                                                                          small (20 to 30 acres [8 to 12 ha]),
their sheep to discourage predators.        well they are tended by the livestock
                                                                                          level, short-grass pastures. Three to
Where effects have been measured,           producer.
                                                                                          four units should be used in larger
however, no difference in losses was
                                            Strobe Lights and Sirens. The                 (40 to 100 acres [16 to 40 ha]), hilly,
detected.
                                            USDA’s Denver Wildlife Research               tall grass, or wooded pastures.

                                                                                                                                     C-59
5. Don’t use EGs in pastures larger        compound that, when applied to              Aversive conditioning is well docu-
          than about 100 acres (40 ha) because    sheep, would prevent coyotes from           mented for averting rodents from food
          their effective range is limited. The   killing them. Tests were conducted          sources, but significant problems must
          device could be useful in larger pas-   with various prey species including         be overcome before the method can be
          tures when placed near areas where      rabbits, chickens, and sheep. Some          used to reduce coyote predation on
          sheep congregate and bed at night.      repellents were applied by dipping tar-     sheep. Coyotes must be induced to eat
                                                  get animals in them, others were            sheeplike baits that have been treated
       6. EGs should be placed on high spots,
                                                  sprayed on, and some were applied in        with the aversive chemical. The chemi-
          where kills have been found, at the
                                                  neck collars or ear tags.                   cal must cause sufficient discomfort,
          edge of wooded areas, near or on
                                                                                              such as vomiting, to cause coyotes to
          bedgrounds, or near suspected           Coyotes rely heavily on visual cues
                                                                                              avoid other baits. Furthermore, the
          coyote travelways. They should be       while stalking, chasing, and killing
                                                                                              avoidance must be transferred to live
          moved to different locations every      their prey. Taste and smell are of lesser
                                                                                              sheep and must persist long enough
          10 to 14 days to reduce the likeli-     importance in actually making the kill.
                                                                                              without reinforcement for the method
          hood of coyotes getting used to         These factors may in part account for
                                                                                              to offer realistic protection to sheep.
          them.                                   the fact that the repellent compounds
                                                  were not able to consistently prevent       To date, pen and field tests with
       To use the EG in open range (herded
                                                  coyotes from killing, although some of      aversive conditioning have yielded
       or range sheep):
                                                  the repellents were obviously offensive     conflicting and inconclusive results. It
       1. The number of EGs used will             to coyotes and prevented them from          does not appear that aversive condi-
          depend on the number of sheep in        consuming the killed prey. Several          tioning is effective in reducing preda-
          the band and the size of the            compounds were tested on sheep              tion, but additional field tests would
          bedground. Four units should be         under field conditions, but none            be useful.
          used to protect bands of 1,000 ewes     appeared to offer significant, pro-
                                                                                              Guarding Animals.
          and their lambs.                        longed protection.
                                                                                              Livestock Guarding Dogs. A live-
       2. When possible, place one EG in the      If an effective chemical repellent were     stock guarding dog is one that gener-
          center of the bedground and the         to be found, the obstacles in bringing it   ally stays with sheep or cattle without
          other three around the edge of the      to industry use would be significant.       harming them and aggressively repels
          bedground. Try to place the units       The compound would not only need            predators. Its protective behaviors are
          on coyote travelways.                   to be effective, but also persistent        largely instinctive, but proper rearing
                                                  enough to withstand weathering while        plays a part. Breeds most commonly
       3. EGs should be placed on high
                                                  posing no undue risk to the sheep,          used today include the Great Pyrenees,
          points, ridge tops, edges of clear-
                                                  other animals, or the environment. It       Komondor, Anatolian Shepherd, and
          ings, or on high rocks or
                                                  would also have to withstand the rig-       Akbash Dog (Fig. 7). Other Old World
          outcroppings. Hang the devices on
                                                  orous Environmental Protection              breeds used to a lesser degree include
          tree limbs 5 to 7 feet (1.5 to 2.1 m)
                                                  Agency (EPA) approval process.              Maremma, Sharplaninetz, and Kuvasz.
          above ground level. If used above
                                                                                              Crossbreeds are also used.
          timberline or in treeless areas, hang   High-frequency sound has also been
          them from a tripod of poles.            tested as a repellent for coyotes, but      The characteristics of each sheep
                                                  the results were no more encouraging        operation will dictate the number of
       4. Herders who bed their sheep tightly
                                                  than for chemical repellents. Coyotes,      dogs required for effective protection
          will have better results than those
                                                  like dogs, responded to particular          from predators. If predators are scarce,
          who allow sheep to bed over large
                                                  sound frequencies and showed some           one dog is sufficient for most fenced
          areas. Sheep that are bedded about
                                                  aversion to sounds broadcast within         pasture operations. Range operations
          200 yards (166 m) or less in diam-
                                                  one foot (30 cm) of their ear. Research-    often use two dogs per band of sheep.
          eter, or are spread out not more
                                                  ers, however, were unable to broad-         The performance of individual dogs
          than 200 to 400 yards (166 to 332 m)
                                                  cast the sound a sufficient distance to     will differ based on age and experi-
          along a ridge top, can usually be
                                                  test the effects under field conditions.    ence. The size, topography, and habitat
          protected with EGs.
                                                                                              of the pasture or range must also be
                                                  Aversive Conditioning. The objec-
       Repellents. The notion of repelling                                                    considered. Relatively flat, open areas
                                                  tive of aversive conditioning is to feed
       coyotes from sheep or calves is                                                        can be adequately covered by one dog.
                                                  a coyote a preylike bait laced with an
       appealing, and during the 1970s, uni-                                                  When brush, timber, ravines, and hills
                                                  aversive agent that causes the coyote
       versity and government researchers                                                     are in the pasture, several dogs may be
                                                  to become ill, resulting in subsequent
       tested a wide variety of potentially re-                                               required, particularly if the sheep are
                                                  avoidance of the prey. Most of the
       pellent chemical compounds on sheep.                                                   scattered. Sheep that flock and form a
                                                  research on this technique has
       Both olfactory (smell) and gustatory                                                   cohesive unit, especially at night, can
                                                  involved the use of lithium chloride, a
       (taste) repellents were examined. The                                                  be protected by one dog more effec-
                                                  salt, as the aversive agent.
       underlying objective was to find a                                                     tively than sheep that are continually

C-60
working unless the dog is chained or
                                                                                           confined while the control takes place.
                                                                                           Dogs caught in a steel trap set for
                                                                                           predators are rarely injured seriously if
                                                                                           they are found and released within a
                                                                                           reasonable period of time. If snares
                                                                                           and traps are used where dogs are
                                                                                           working, the producer should: (1)
                                                                                           encourage the use of sets and devices
                                                                                           that are likely not to injure the dog if it
                                                                                           is caught, and (2) know where traps
                                                                                           and snares are set so they can be
                                                                                           checked if a dog is missing. Aerial
                                                                                           hunting, as well as calling and shoot-
                                                                                           ing coyotes, should pose no threat to
                                                                                           guarding dogs. Ensuring the safety of
                                                                                           the dog is largely the producer’s
                                                                                           responsibility.
                                                                                           Dogs may be viewed as a first line of
Fig. 7. Livestock guarding dog (Akbash dog)                                                defense against predation in sheep and
                                                                                           cow/calf operations in some cases.
                                                                                           Their effectiveness can be enhanced by
scattered and bedded in a number of           to the possibility that the dog may          good livestock management and by
locations.                                    roam onto their property and that            eliminating predators with suitable
                                              some predator control devices such as        removal techniques.
The goal with a new puppy is to chan-
                                              traps, snares, and M-44s present a
nel its natural instincts to produce a                                                     Donkeys. Although the research has
                                              danger to it. Many counties enforce
mature guardian dog with the desired                                                       not focused on donkeys as it has on
                                              stringent laws regarding owner
characteristics. This is best accom-                                                       guarding dogs, they are gaining in
                                              responsibility for damage done by
plished by early and continued asso-                                                       popularity as protectors of sheep and
                                              roaming dogs. It is in the best interests
ciation with sheep to produce a bond                                                       goat flocks in the United States. A
                                              of the owner, dog, and community to
between the dog and sheep. The opti-                                                       recent survey showed that in Texas
                                              train the dog to stay in its designated
mum time to acquire a pup is between                                                       alone, over 2,400 of the 11,000 sheep
                                              area.
7 and 8 weeks of age. The pup should                                                       and goat producers had used donkeys
be separated from litter mates and            The use of guarding dogs does not            as guardians.
placed with sheep, preferably lambs,          eliminate the need for other predation       The terms donkey and burro are syn-
in a pen or corral from which it can’t        control actions. They should, however,       onymous (the Spanish translation of
escape. This socialization period             be compatible with the dog’s behavior.       donkey is burro) and are used inter-
should continue with daily checks             Toxicants (including some insecticides       changeably. Donkeys are generally
from the producer until the pup is            and rodenticides) used to control vari-      docile to people, but they seem to
about 16 weeks old. Daily checks don’t        ous pest species can be extremely haz-       have an inherent dislike of dogs and
necessarily include petting the pup.          ardous to dogs and are therefore not         other canids, including coyotes and
The primary bond should be between            compatible with the use of guarding          foxes. The typical response of a don-
the dog and the sheep, not between the        dogs.                                        key to an intruding canid may include
dog and humans. The owner, how-                                                            braying, bared teeth, a running attack,
                                              The M-44 is particularly hazardous to
ever, should be able to catch and han-                                                     kicking, and biting. Most likely it is
                                              dogs. Some people have successfully
dle the dog to administer health care                                                      acting out of aggression toward the
                                              trained their dogs to avoid M-44s by
or to manage the livestock. At about 4                                                     intruder rather than to protect the
                                              allowing the dog to set off an M-44
months, the pup can be released into a                                                     sheep. There is little information on a
                                              filled with pepper or by rigging the
larger pasture to mingle with the other                                                    donkey’s effectiveness with noncanid
                                              device to a rat trap. The unpleasant
sheep.                                                                                     predators such as bears, mountain
                                              experience may teach the dog to avoid
A guarding dog will likely include            M-44s, but the method is not fool-           lions, bobcats, or birds of prey.
peripheral areas in its patrolling. Some      proof—one error by the dog, and the          Reported success of donkeys in reduc-
have been known to chase vehicles             result is usually fatal. With the excep-     ing predation is highly variable.
and wildlife and threaten children and        tion of toxic collars, which are not legal   Improper husbandry or rearing prac-
cyclists. These activities should be dis-     in all states, toxicants should not be       tices and unrealistic expectations
couraged. Neighbors should be alerted         used in areas where guarding dogs are

                                                                                                                                         C-61
probably account for many failures.        Llamas. Like donkeys, llamas have an         Toxicants
       Donkeys are significantly cheaper to       inherent dislike of canids, and a grow-
                                                                                               Pesticides have historically been an
       obtain and care for than guarding          ing number of livestock producers are
                                                                                               important component in an integrated
       dogs, and they are probably less prone     successfully using llamas to protect
                                                                                               approach to controlling coyote dam-
       to accidental death and premature          their sheep. A recent study of 145
                                                                                               age, but their use is extremely
       mortality than dogs. They may pro-         ranches where guard llamas were used
                                                                                               restricted today by federal and state
       vide a longer period of useful life than   to protect sheep revealed that average
                                                                                               laws. All pesticides used in the United
       a guarding dog, and they can be used       losses of sheep to predators decreased
                                                                                               States must be registered with the EPA
       with relative safety in conjunction with   from 26 to 8 per year after llamas were
                                                                                               under the provisions of FIFRA and
       snares, traps, M-44s, and toxic collars.   employed. Eighty percent of the
                                                                                               must be used in accordance with label
                                                  ranchers surveyed were “very satis-
       Researchers and livestock producers                                                     directions. Increasingly restrictive
                                                  fied” or “satisfied” with their llamas.
       have identified several key points to                                                   regulations implemented by EPA
                                                  Llamas reportedly bond with sheep
       consider when using a donkey for pre-                                                   under the authority of FIFRA, the
                                                  within hours and offer advantages
       dation control:                                                                         National Environmental Policy Act
                                                  over guarding dogs similar to those
                                                                                               (NEPA), presidential order, and the
       1. Use only a jenny or a gelded jack.      described for donkeys.
                                                                                               Endangered Species Act have resulted
          Intact jacks are too aggressive and
                                                  Other Animals. USDA’s Agricultural           in the near elimination of toxicants
          may injure livestock. Some jennies
                                                  Research Service tested the bonding of       legally available for predator damage
          and geldings may also injure live-
                                                  sheep to cattle as a method of protect-      control.
          stock. Select donkeys from
                                                  ing sheep from coyote predation.
          medium-sized stock.                                                                  The only toxicants currently registered
                                                  There was clearly some protection
                                                                                               for mammalian predator damage con-
       2. Use only one donkey per group of        afforded the sheep that remained near
                                                                                               trol are sodium cyanide, used in the
          sheep. The exception may be a           cattle. Whether this protection resulted
                                                                                               M-44 ejector device, and Compound
          jenny with a foal. When two or          from direct action by the cattle or by
                                                                                               1080 (sodium monofluoroacetate), for
          more adult donkeys are together or      the coyotes’ response to a novel stimu-
                                                                                               use in the livestock protection collar.
          with a horse, they usually stay         lus is uncertain. Later studies with
                                                                                               These toxicants are Restricted Use Pes-
          together, not necessarily near the      goats, sheep, and cattle confirmed that
                                                                                               ticides and may be used only by certi-
          sheep. Also avoid using donkeys in      when either goats or sheep remained
                                                                                               fied pesticide applicators. Information
          adjacent pastures since they may        near cattle, they were protected from
                                                                                               on registration status and availability
          socialize across the fence and ignore   predation by coyotes. Conversely,
                                                                                               of these products in individual states
          the sheep.                              goats or sheep that grazed apart from
                                                                                               may be obtained from the respective
                                                  cattle, even those that were bonded,
       3. Allow about 4 to 6 weeks for a                                                       state’s department of agriculture.
                                                  were readily preyed on by coyotes.
          naive donkey to bond to the sheep.
                                                                                               Sodium Cyanide in the M-44. The
          Stronger bonding may occur when         There are currently no research data
                                                                                               M-44 is a spring-activated device used
          a donkey is raised from birth with      available on the ideal ratio of cattle to
                                                                                               to expel sodium cyanide into an
          sheep.                                  sheep, the breeds of cattle, age of cattle
                                                                                               animal’s mouth. It is currently regis-
                                                  most likely to be used successfully, or
       4. Avoid feeds or supplements con-                                                      tered by EPA for use by trained per-
                                                  on the size of bonded groups to obtain
          taining monensin or lasolacid. They                                                  sonnel in the control of depredating
                                                  maximum protection from predation.
          are poisonous to donkeys.                                                            coyotes, foxes, and dogs.
                                                  Multispecies grazing offers many
       5. Remove the donkey during lamb-          advantages for optimum utilization of        The M-44 consists of a capsule holder
          ing, particularly if lambing in con-    forage, and though additional study          wrapped in an absorbent material, an
          finement, to avoid injuries to lambs    and experience is needed, it may also        ejector mechanism, a capsule contain-
          or disruption of the lamb-ewe bond.     be a tool for coyote damage control.         ing approximately 0.9 grams of a pow-
                                                                                               dered sodium cyanide mixture, and a
       6. Test a new donkey’s response to         Any animal that displays aggressive
                                                                                               5- to 7-inch (15- to 18-cm) hollow stake
          canids by challenging it with a dog     behavior toward intruding coyotes
                                                                                               (Fig. 8). For most effective use, set
          in a pen or small pasture. Discard      may offer some benefit in deterring
                                                                                               M-44s in locations similar to those for
          donkeys that don’t show overt           predation. Other types of animals
                                                                                               good trap sets. Drive the hollow stake
          aggression to an intruding dog.         reportedly used for predation control
                                                                                               into the ground. Cock the ejector unit
                                                  include goats, mules, and ostriches.
       7. Use donkeys in smaller (less than                                                    and secure it in the stake. Screw the
                                                  Coyotes in particular are suspicious of
          600 acres [240 ha]), relatively open                                                 wrapped capsule holder containing the
                                                  novel stimuli. This behavior is most
          pastures with not more than 200 to                                                   cyanide capsule onto the ejector unit,
                                                  likely the primary reason that many
          300 head of livestock. Large pas-                                                    and apply fetid meat bait to the cap-
                                                  frightening tactics show at least tem-
          tures with rough terrain and vegeta-                                                 sule holder. Coyotes attracted by the
                                                  porary effectiveness.
          tion and widely scattered livestock                                                  bait will try to bite the baited capsule
          lessen the effectiveness of a donkey.                                                holder. When the M-44 is pulled, the

C-62
their ability to remain effective during   The LP collar must be used in conjunc-
                                                    rain, snow, and freezing conditions.       tion with specific sheep and goat hus-
           c                                                                                   bandry practices to be most effective.
                                                    While M-44s can be used effectively as
                                                                                               Coyote attacks must be directed or tar-
                                                    part of an integrated damage control
                                                                                               geted at collared livestock. This may
                                  d                 program, they do have several disad-
                                                                                               be accomplished by temporarily plac-
                                                    vantages. Because canids are less
                                                                                               ing a “target” flock of perhaps 20 to 50
                                                    responsive to food-type baits during
                                                                                               collared lambs or kids and their
                                                    warm weather when natural foods are
                                                                                               uncollared mothers in a pasture where
                                                    usually abundant, M-44s are not as
                                                                                               coyote predation is likely to occur,
                                                    effective during warmer months as
                                                                                               while removing other sheep or goats
                                                    they are in cooler weather. M-44s are
                                                                                               from that vicinity. In situations where
                                                    subject to a variety of mechanical mal-
                                                                                               LP collars have been used and found
                                                    functions, but these problems can be
                                                                                               ineffective, the common cause of fail-
                b                                   minimized if a regular maintenance
                                                                                               ure has been poor or ineffective target-
                                                    schedule is followed. A further disad-
                                                                                               ing. It is difficult to ensure effective
                                                    vantage is the tendency for the cyanide
                                                                                               targeting if depredations are occurring
                                                    in the capsules to absorb moisture over
                                                                                               infrequently. In most instances, only a
                                                    time and to cake, becoming ineffective.
                                                                                               high and regular frequency of depre-
                                                    Maximum effectiveness of M-44s is
                                                                                               dations will justify spending the time,
                                                    hampered by the requirement to fol-
                                                                                               effort, and money necessary to become
                                                    low 26 use restrictions established by
                                                                                               trained and certified, purchase collars,
                                                    the EPA in the interest of human and
                                                                                               and use them properly.
                                                    environmental safety. The M-44 is not
                                                    registered for use in all states, and in   The outstanding advantage in using
                                                    those where it is registered, the state    the LP collar is its selectivity in elimi-
                                                    may impose additional use restric-         nating individual coyotes that are
                                                    tions. A formal training program is        responsible for killing livestock. The
                                                    required before use of M-44s. Some         collar may also be useful in removing
                                                    states allow its use only by federal       depredating coyotes that have eluded
           a                                        ADC specialists, whereas other states      other means of control. Disadvantages
                                                    may allow M-44s to be used by trained      include the cost of collars (approxi-
                                                    and certified livestock producers.         mately $20 each) and livestock that
                                                                                               must be sacrificed, more intensive
Fig. 8. The M-44 device consists of the (a) base,   1080 Livestock Protection Collar.
(b) ejector, (c) capsule holder, and (d) cyanide-                                              management practices, and the costs
                                                    The livestock protection collar (LP
containing plastic capsule.                                                                    and inconvenience of complying with
                                                    collar or toxic collar) is a relatively
                                                                                               use restrictions, including require-
                                                    new tool used to selectively kill
                                                                                               ments for training, certification, and
spring-activated plunger propels                    coyotes that attack sheep or goats.
                                                                                               record keeping. One use restriction
sodium cyanide into the animal’s                    Collars are placed on sheep or goats
                                                                                               limits the collars to use in fenced pas-
mouth, resulting in death within a few              that are pastured where coyotes are
                                                                                               tures only. They cannot be used to pro-
seconds.                                            likely to attack. Each collar contains a
                                                                                               tect sheep on open range. Also, collars
                                                    small quantity (300 mg) of Compound
The M-44 is very selective for canids                                                          are not widely available, because they
                                                    1080 solution. The collars do not
because of the attractants used and the                                                        are registered for use in only a few
                                                    attract coyotes, but because of their
unique requirement that the device be                                                          states.
                                                    design and position on the throat,
triggered by pulling on it. While the               most attacking coyotes will puncture
use of traps or snares may present a                                                           Fumigants
                                                    the collar and ingest a lethal amount of
hazard to livestock, M-44s can be used              the toxicant. Unlike sodium cyanide,       Carbon monoxide is an effective bur-
with relative safety in pastures where              1080 is slow-acting, and a coyote          row fumigant recently re-registered by
livestock are present. Although not                 ingesting the toxicant will not exhibit    the EPA. Gas cartridges, which contain
recommended, they can also be used                  symptoms or die for several hours. As      65% sodium nitrate and 35% charcoal,
in the presence of livestock guarding               a result, sheep or goats that are          produce carbon monoxide, carbon
dogs if the dogs are first successfully             attacked are usually killed. The collar    dioxide, and other noxious gases when
conditioned to avoid the devices. This              is registered only for use against         ignited. They were registered by the
can be done by allowing them to pull                coyotes and may be placed only on          EPA in 1981 for control of coyotes in
an M-44 loaded with pepper. An addi-                sheep or goats.                            dens only. This is the only fumigant
tional advantage of M-44s over traps is                                                        currently registered for this purpose.

                                                                                                                                            C-63
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